About the Authors

Morgan N Price

Affiliations Lawrence Berkeley National Laboratory, Berkeley, California, United States of America , Virtual Institute for Microbial Stress and Survival, University of California San Francisco, San Francisco, California, United States of America

Adam P Arkin

Affiliations Lawrence Berkeley National Laboratory, Berkeley, California, United States of America , Virtual Institute for Microbial Stress and Survival, University of California San Francisco, San Francisco, California, United States of America , Department of Bioengineering, University of California Berkeley, Berkeley, California, United States of America , Howard Hughes Medical Institute, University of California Berkeley, Berkeley, California, United States of America

Eric J Alm

To whom correspondence should be addressed. E-mail: ejalm@mit.edu

¤ Current address: Department of Civil and Environmental Engineering and Division of Biological Engineering, Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Affiliations Lawrence Berkeley National Laboratory, Berkeley, California, United States of America , Virtual Institute for Microbial Stress and Survival, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MNP and EJA conceived and designed the experiments. MNP performed the experiments. MNP analyzed the data. APA supervised the work. MNP, APA, and EJA wrote the paper.