About the Authors

María José Aranzana

Contributed equally to this work with: María José Aranzana, Sung Kim, Keyan Zhao

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Sung Kim

Contributed equally to this work with: María José Aranzana, Sung Kim, Keyan Zhao

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Keyan Zhao

Contributed equally to this work with: María José Aranzana, Sung Kim, Keyan Zhao

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Erica Bakker

Affiliation Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Matthew Horton

Affiliation Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Katrin Jakob

Affiliation Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Clare Lister

Affiliation Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

John Molitor

Affiliation Department of Preventive Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Chikako Shindo

Affiliation Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Chunlao Tang

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Christopher Toomajian

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Brian Traw

Affiliation Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Honggang Zheng

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Joy Bergelson

Affiliation Department of Ecology and Evolution, University of Chicago, Chicago, Illinois, United States of America

Caroline Dean

Affiliation Cell and Developmental Biology, John Innes Centre, Norwich, United Kingdom

Paul Marjoram

Affiliation Department of Preventive Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Magnus Nordborg

To whom correspondence should be addressed. E-mail: magnus@usc.edu

Affiliation Molecular and Computational Biology, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JB, CD, PM, and MN designed and directed the overall project. MJA, CL, CS, CT, HZ, CD, and MN generated the flowering time data. EB, MH, KJ, BT, and JB generated the resistance data. MJA, CT, CT, BT, and HJ generated the candidate locus polymorphism data. MJA, SK, KZ, JM, PM, and MN analyzed the data. SK, KZ, and MN wrote the paper.