About the Authors

Naomi V. Bradbury

N.Bradbury@warwick.ac.uk

Affiliations School of Veterinary Medicine and Science, University of Nottingham, Leicestershire, United Kingdom, School of Life Sciences, University of Warwick, Coventry, United Kingdom

William J. M. Probert

Affiliations School of Veterinary Medicine and Science, University of Nottingham, Leicestershire, United Kingdom, School of Life Sciences, University of Warwick, Coventry, United Kingdom, Mathematics Institute, University of Warwick, Coventry, United Kingdom

Katriona Shea

Affiliations Center for Infectious Disease Dynamics, Department of Biology, Eberly College of Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Department of Biology and Intercollege Graduate Degree Program in Ecology, 208 Mueller Laboratory, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Michael C. Runge

Affiliation US Geological Survey, Patuxent Wildlife Research Center, Laurel, Maryland, United States of America

Christopher J. Fonnesbeck

Affiliation Department of Biostatistics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Matt J. Keeling

Affiliations School of Life Sciences, University of Warwick, Coventry, United Kingdom, Mathematics Institute, University of Warwick, Coventry, United Kingdom

Matthew J. Ferrari

Affiliations Center for Infectious Disease Dynamics, Department of Biology, Eberly College of Science, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Department of Biology and Intercollege Graduate Degree Program in Ecology, 208 Mueller Laboratory, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Michael J. Tildesley

Affiliations School of Veterinary Medicine and Science, University of Nottingham, Leicestershire, United Kingdom, School of Life Sciences, University of Warwick, Coventry, United Kingdom, Mathematics Institute, University of Warwick, Coventry, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NVB WJMP KS MCR CJF MJK MJF MJT. Performed the experiments: MJT. Analyzed the data: NVB WJMP MJT. Wrote the paper: NVB WJMP KS MCR CJF MJK MJF MJT.