About the Authors

Rory M. Donovan

Affiliations Joint CMU-Pitt Ph.D. Program in Computational Biology, Pittsburgh, Pennsylvania, United States of America, Department of Computational and Systems Biology, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Jose-Juan Tapia

Affiliations Joint CMU-Pitt Ph.D. Program in Computational Biology, Pittsburgh, Pennsylvania, United States of America, Department of Computational and Systems Biology, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Devin P. Sullivan

Affiliations Joint CMU-Pitt Ph.D. Program in Computational Biology, Pittsburgh, Pennsylvania, United States of America, Computational Biology Department, School of Computer Science, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

James R. Faeder

Affiliation Department of Computational and Systems Biology, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Robert F. Murphy

Affiliation Computational Biology Department, School of Computer Science, Carnegie Mellon University, Pittsburgh, Pennsylvania, United States of America

Markus Dittrich

Affiliations Department of Computational and Systems Biology, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Pittsburgh Supercomputing Center, Pittsburgh, Pennsylvania, United States of America

Daniel M. Zuckerman

ddmmzz@pitt.edu

Affiliation Department of Computational and Systems Biology, School of Medicine, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RMD JRF RFM MD DMZ. Performed the experiments: RMD JJT DPS. Analyzed the data: RMD DMZ. Wrote the paper: RMD JJT DPS JRF RFM MD DMZ.