About the Authors

Mayu Sugiyama

Contributed equally to this work with: Mayu Sugiyama, Takashi Saitou

Affiliation Laboratory for Cell Function and Dynamics, Advanced Technology Development Group, Brain Science Institute, RIKEN, Wako-city, Saitama, Japan

Takashi Saitou

Contributed equally to this work with: Mayu Sugiyama, Takashi Saitou

Affiliation Department of Molecular Medicine for Pathogenesis, Graduate School of Medicine, Ehime University, Shitsukawa, Toon-city, Ehime, Japan

Hiroshi Kurokawa

Affiliation Laboratory for Cell Function and Dynamics, Advanced Technology Development Group, Brain Science Institute, RIKEN, Wako-city, Saitama, Japan

Asako Sakaue-Sawano

Affiliation Laboratory for Cell Function and Dynamics, Advanced Technology Development Group, Brain Science Institute, RIKEN, Wako-city, Saitama, Japan

Takeshi Imamura

timamura@m.ehime-u.ac.jp (TIm); matsushi@brain.riken.go.jp (AM); iimura@m.ehime-u.ac.jp (TIi)

Affiliations Department of Molecular Medicine for Pathogenesis, Graduate School of Medicine, Ehime University, Shitsukawa, Toon-city, Ehime, Japan, Division of Bio-Imaging, Proteo-Science Center (PROS), Ehime University, Shitsukawa, Toon-city, Ehime, Japan, Translational Research Center, Ehime University Hospital, Shitsukawa, Toon-city, Ehime, Japan

Atsushi Miyawaki

timamura@m.ehime-u.ac.jp (TIm); matsushi@brain.riken.go.jp (AM); iimura@m.ehime-u.ac.jp (TIi)

Affiliation Laboratory for Cell Function and Dynamics, Advanced Technology Development Group, Brain Science Institute, RIKEN, Wako-city, Saitama, Japan

Tadahiro Iimura

timamura@m.ehime-u.ac.jp (TIm); matsushi@brain.riken.go.jp (AM); iimura@m.ehime-u.ac.jp (TIi)

Affiliations Laboratory for Cell Function and Dynamics, Advanced Technology Development Group, Brain Science Institute, RIKEN, Wako-city, Saitama, Japan, Division of Bio-Imaging, Proteo-Science Center (PROS), Ehime University, Shitsukawa, Toon-city, Ehime, Japan, Translational Research Center, Ehime University Hospital, Shitsukawa, Toon-city, Ehime, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AM TIi. Performed the experiments: MS TS TIi. Analyzed the data: MS TS TIi. Contributed reagents/materials/analysis tools: HK ASS TIm. Wrote the paper: TS TIi.