About the Authors

Sunhwan Jo

Affiliation Department of Molecular Biosciences and Center for Bioinformatics, The University of Kansas, Lawrence, Kansas, United States of America

Hui Sun Lee

Affiliation Department of Molecular Biosciences and Center for Bioinformatics, The University of Kansas, Lawrence, Kansas, United States of America

Jeffrey Skolnick

Affiliation Center for the Study of Systems Biology, School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Wonpil Im

wonpil@ku.edu

Affiliation Department of Molecular Biosciences and Center for Bioinformatics, The University of Kansas, Lawrence, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJ WI. Performed the experiments: SJ HSL. Analyzed the data: SJ HSL JS WI. Contributed reagents/materials/analysis tools: SJ HSL. Wrote the paper: SJ HSL JS WI.