About the Authors

Alexander Dilthey

Contributed equally to this work with: Alexander Dilthey, Stephen Leslie

dilthey@well.ox.ac.uk

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, Oxford, United Kingdom

Stephen Leslie

Contributed equally to this work with: Alexander Dilthey, Stephen Leslie

Affiliation Murdoch Childrens Research Institute, Royal Children's Hospital, Parkville, Victoria, Australia

Loukas Moutsianas

Affiliation Wellcome Trust Centre for Human Genetics, Oxford, United Kingdom

Judong Shen

Affiliation Quantitative Sciences, GlaxoSmithKline, Research Triangle Park, North Carolina, United States of America

Charles Cox

Affiliation Quantitative Sciences, GlaxoSmithKline, Stevenage, United Kingdom

Matthew R. Nelson

Affiliation Quantitative Sciences, GlaxoSmithKline, Research Triangle Park, North Carolina, United States of America

Gil McVean

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AD SL MRN GM. Performed the experiments: AD SL LM JS. Analyzed the data: AD SL. Contributed reagents/materials/analysis tools: CC. Wrote the paper: AD SL MRN GM.