About the Authors

David M. Richards

Affiliation Computational and Systems Biology, John Innes Centre, Norwich Research Park, Norwich, Norfolk, United Kingdom

Emma Greer

Affiliation Cell & Developmental Biology, John Innes Centre, Norwich Research Park, Norwich, Norfolk, United Kingdom

Azahara C. Martin

Affiliation Cell & Developmental Biology, John Innes Centre, Norwich Research Park, Norwich, Norfolk, United Kingdom

Graham Moore

Affiliation Crop Genetics, John Innes Centre, Norwich Research Park, Norwich, Norfolk, United Kingdom

Peter J. Shaw

peter.shaw@jic.ac.uk (PJS); martin.howard@jic.ac.uk (MH)

Affiliation Cell & Developmental Biology, John Innes Centre, Norwich Research Park, Norwich, Norfolk, United Kingdom

Martin Howard

peter.shaw@jic.ac.uk (PJS); martin.howard@jic.ac.uk (MH)

Affiliation Computational and Systems Biology, John Innes Centre, Norwich Research Park, Norwich, Norfolk, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DMR EG GM PJS MH. Performed the experiments: EG ACM. Analyzed the data: DMR MH. Wrote the paper: DMR EG ACM GM PJS MH. Designed and performed the computational modelling: DMR MH.