About the Authors

Haipeng Xing

Contributed equally to this work with: Haipeng Xing, Yifan Mo, Will Liao

xing@ams.sunysb.edu

Affiliation Applied Mathematics and Statistics, Stony Brook University, Stony Brook, New York, United States of America

Yifan Mo

Contributed equally to this work with: Haipeng Xing, Yifan Mo, Will Liao

Affiliations Applied Mathematics and Statistics, Stony Brook University, Stony Brook, New York, United States of America, Computational Biology, Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Will Liao

Contributed equally to this work with: Haipeng Xing, Yifan Mo, Will Liao

Affiliations Applied Mathematics and Statistics, Stony Brook University, Stony Brook, New York, United States of America, Computational Biology, Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America

Michael Q. Zhang

Address Current address: Department of Molecular and Cell Biology, The University of Texas at Dallas, Richardson, Texas, United States of America.

Affiliations Computational Biology, Cold Spring Harbor Laboratory, Cold Spring Harbor, New York, United States of America, Bioinformatics Division, Tsinghua University, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HX YM WL MQZ. Analyzed the data: HX YM WL. Contributed reagents/materials/analysis tools: HX MQZ. Wrote the paper: HX YM WL.