About the Authors

T. Déirdre Hollingsworth, Roy M. Anderson
MRC Centre for Outbreak Control and Analysis, Department of Infectious Disease Epidemiology, Imperial College London, London, United Kingdom

Don Klinkenberg, Hans Heesterbeek
Theoretical Epidemiology, Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Corresponding Author


Competing Interests

Roy M. Anderson is a non-executive member of the board of GlaxoSmithKline (GSK).

Author Contributions

Conceived and designed the experiments: TDH DK HH RMA. Performed the experiments: TDH DK. Analyzed the data: TDH DK. Wrote the paper: TDH DK HH RMA.