About the Authors

Ron Skupsky

Affiliation California Institute for Quantitative Biosciences, University of California, Berkeley, Berkeley, California, United States of America

John C. Burnett

Address Current address: Department of Molecular and Cell Biology, Beckman Research Institute of the City of Hope, Duarte, California, United States of America.

Affiliations Department of Chemical Engineering, University of California, Berkeley, Berkeley, California, United States of America, Helen Wills Neuroscience Institute, University of California, Berkeley, Berkeley, California, United States of America

Jonathan E. Foley

Affiliation UCB/UCSF Joint-Graduate-Group-in-Bioengineering, University of California, Berkeley, Berkeley, California, United States of America

David V. Schaffer

schaffer@berkeley.edu (DVS); aparkin@lbl.gov (APA)

Affiliations California Institute for Quantitative Biosciences, University of California, Berkeley, Berkeley, California, United States of America, Department of Chemical Engineering, University of California, Berkeley, Berkeley, California, United States of America, Helen Wills Neuroscience Institute, University of California, Berkeley, Berkeley, California, United States of America, Department of Bioengineering, University of California, Berkeley, Berkeley, California, United States of America

Adam P. Arkin

schaffer@berkeley.edu (DVS); aparkin@lbl.gov (APA)

Affiliations California Institute for Quantitative Biosciences, University of California, Berkeley, Berkeley, California, United States of America, Department of Bioengineering, University of California, Berkeley, Berkeley, California, United States of America, Physical Biosciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RS JCB DVS APA. Performed the experiments: JCB JEF. Analyzed the data: RS. Contributed reagents/materials/analysis tools: RS JCB JEF DVS. Wrote the paper: RS JCB JEF DVS APA.