About the Authors

Philippe Lemey
Department of Microbiology and Immunology, Katholieke Universiteit Leuven, Leuven, Belgium

Andrew Rambaut
Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, United Kingdom

Alexei J. Drummond
Department of Computer Science, University of Auckland, Auckland, New Zealand

Marc A. Suchard
Departments of Biomathematics and Human Genetics, David Geffen School of Medicine, University of California, Los Angeles, California, United States of America

Marc A. Suchard
Department of Biostatistics, School of Public Health, University of California, Los Angeles, California, United States of America

Corresponding Author

philippe.lemey@uz.kuleuven.be

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: PL. Wrote the paper: PL MAS. Designed the study: PL MAS. Provided programming assistance: AR AJD. Conceived the original idea and developed the software implementation: MAS.