About the Authors

Shigeru Shinomoto, Hideaki Kim, Takeaki Shimokawa, Nanae Matsuno
Graduate School of Science, Kyoto University, Sakyo-ku, Kyoto, Japan

Shintaro Funahashi
Kokoro Research Center, Kyoto University, Sakyo-ku, Kyoto, Japan

Keisetsu Shima
Department of Physiology, Tohoku University School of Medicine, Aoba-ku, Sendai, Japan

Ichiro Fujita, Hiroshi Tamura, Taijiro Doi
Laboratory for Cognitive Neuroscience, Graduate School of Frontier Biosciences, Osaka University, Toyonaka, Osaka, Japan

Kenji Kawano, Naoko Inaba, Tadashi Ogawa
Department of Integrative Brain Science, Graduate School of Medicine, Kyoto University, Sakyo-ku, Kyoto, Japan

Kikuro Fukushima, Sergei Kurkin
Department of Physiology, Hokkaido University School of Medicine, Sapporo, Japan

Kiyoshi Kurata
Department of Physiology, Hirosaki University School of Medicine, Hirosaki, Japan

Masato Taira
Division of Applied System Neuroscience, Advanced Medical Research Center, Nihon University Graduate School of Medical Science, Tokyo, Japan

Ken-Ichiro Tsutsui
Division of Systems Neuroscience, Tohoku University Graduate School of Life Sciences, Aoba-ku, Sendai, Japan

Hidehiko Komatsu, Tadashi Ogawa, Kowa Koida
Division of Sensory and Cognitive Information, National Institute for Physiological Sciences, Myodaiji, Okazaki, Aichi, Japan

Jun Tanji
Tamagawa University Brain Science Institute, Machida, Tokyo, Japan

Keisuke Toyama
ATR Computational Neuroscience Laboratories, Seika-cho, Soraku-gun, Kyoto, Japan

Corresponding Author

shinomoto@scphys.kyoto-u.ac.jp

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: S. Shinomoto K. Toyama. Performed the experiments: S. Funahashi K. Shima I. Fujita H. Tamura T. Doi K. Kawano N. Inaba K. Fukushima S. Kurkin K. Kurata M. Taira K-I. Tsutsui H. Komatsu T. Ogawa K. Koida J. Tanji. Analyzed the data: S. Shinomoto H. Kim T. Shimokawa N. Matsuno. Contributed reagents/materials/analysis tools: H. Kim T. Shimokawa N. Matsuno. Wrote the paper: S. Shinomoto K. Toyama.