About the Authors

Shobhit Gupta, John A. Stamatoyannopoulos, William Stafford Noble
Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Jonathan Dennis, Robert Kingston
Department of Molecular Biology, Massachusetts General Hospital, Boston, Massachusetts, United States of America

Robert E. Thurman
Division of Medical Genetics, University of Washington, Seattle, Washington, United States of America

William Stafford Noble
Department of Computer Science and Engineering, University of Washington, Seattle, Washington, United States of America

Corresponding Authors

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SG JD JAS WSN. Performed the experiments: SG JD. Analyzed the data: SG JD RET. Wrote the paper: SG JD RK JAS WSN.