About the Authors

Shenghua Li, Paul Brazhnik, John J Tyson
Department of Biological Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Bruno Sobral, John J Tyson
Virginia Bioinformatics Institute, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Corresponding Author

tyson@vt.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SL, PB, and JJT conceived, designed, and performed the research and wrote the paper. BS revised the manuscript for important intellectual content. PB, BS, and JJT obtained funding for this research.