About the Authors

Sergey Kryazhimskiy
Program in Applied and Computational Mathematics, Princeton University, Princeton, New Jersey, United States of America

Ulf Dieckmann
Evolution and Ecology Program, International Institute for Applied Systems Analysis, Laxenburg, Austria

Simon A Levin, Jonathan Dushoff
Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Jonathan Dushoff
Fogarty International Center, National Institute of Health, Bethesda, Maryland, United States of America

Corresponding Author

skryazhi@princeton.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SK, UD, and JD designed the model. SK wrote the code. SK, UD, SAL, and JD wrote the paper.