About the Authors
Department of Biology, Emory University, Atlanta, Georgia, United States of America
Department of Mathematics, University of Florida, Gainesville Florida, United States of America
Emory Vaccine Center, Emory University, Atlanta, Georgia, United States of America
The authors have declared that no competing interests exist.
R. Antia and V. V. Ganusov conceived the basic ideas. R. Ahmed helped formulate the problem and consider the implications for experimental work. R. Antia and V. V. Ganusov derived analytical approximations. V. V. Ganusov performed the simulations. V. V. Ganusov and R. Antia wrote the paper. S. S. Pilyugin contributed to the derivations and helped to modify the text.