About the Authors

Vitaly V Ganusov, Rustom Antia
Department of Biology, Emory University, Atlanta, Georgia, United States of America

Sergei S Pilyugin
Department of Mathematics, University of Florida, Gainesville Florida, United States of America

Rafi Ahmed
Emory Vaccine Center, Emory University, Atlanta, Georgia, United States of America

Corresponding Author

rantia@emory.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

R. Antia and V. V. Ganusov conceived the basic ideas. R. Ahmed helped formulate the problem and consider the implications for experimental work. R. Antia and V. V. Ganusov derived analytical approximations. V. V. Ganusov performed the simulations. V. V. Ganusov and R. Antia wrote the paper. S. S. Pilyugin contributed to the derivations and helped to modify the text.