About the Authors

Jakob Skou Pedersen

To whom correspondence should be addressed. E-mail: jsp@soe.ucsc.edu

Affiliation Center for Biomolecular Science and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Gill Bejerano

Affiliation Center for Biomolecular Science and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Adam Siepel

Address ¤ Current address: Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Affiliation Center for Biomolecular Science and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Kate Rosenbloom

Affiliation Center for Biomolecular Science and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Kerstin Lindblad-Toh

Affiliation Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Eric S Lander

Affiliation Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America

Jim Kent

Affiliation Center for Biomolecular Science and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America

Webb Miller

Affiliation Center for Comparative Genomics and Bioinformatics, Pennsylvania State University, University Park, Pennsylvania, United States of America

David Haussler

Affiliations Center for Biomolecular Science and Engineering, University of California Santa Cruz, Santa Cruz, California, United States of America , Howard Hughes Medical Institute, University of California Santa Cruz, Santa Cruz, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JSP, GB, and DH conceived and designed the experiments. JSP and GB performed the experiments. JSP analyzed the data. JSP, AS, KR, KLT, ESL, JK, and WM contributed reagents/materials/analysis tools. JSP and DH wrote the paper.