About the Authors

Hiroyuki Kurata

Contributed equally to this work with: Hiroyuki Kurata, Hana El-Samad

To whom correspondence should be addressed. E-mail: kurata@bio.kyutech.ac.jp

Affiliation Department of Bioscience and Bioinformatics, Kyushu Institute of Technology, Iizuka, Fukuoka, Japan

Hana El-Samad

Contributed equally to this work with: Hiroyuki Kurata, Hana El-Samad

Affiliations Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America , California Institute for Quantitative Biomedical Research, University of California San Francisco, San Francisco, California, United States of America

Rei Iwasaki

Affiliation Department of Bioscience and Bioinformatics, Kyushu Institute of Technology, Iizuka, Fukuoka, Japan

Hisao Ohtake

Affiliation Department of Biotechnology, Graduate School of Engineering, Osaka University, Osaka, Japan

John C Doyle

Affiliation Control and Dynamical Systems, California Institute of Technology, Pasadena, California, United States of America

Irina Grigorova

Affiliations Department of Cell and Tissue Biology, University of California San Francisco, San Francisco, California, United States of America , Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Carol A Gross

Affiliations Department of Cell and Tissue Biology, University of California San Francisco, San Francisco, California, United States of America , Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Mustafa Khammash

Affiliation Department of Mechanical Engineering, University of California Santa Barbara, Santa Barbara, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HK, HES, HO, JCD, CAG, and MK conceived and designed the experiments. HK and RI performed the experiments. HK, HES, and CAG analyzed the data. HES, JCD, IG, CAG, and MK contributed reagents/materials/analysis tools. HK, HES, JCD, CAG, and MK wrote the paper.