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Contributed equally to this work with:
Kevin M. Wright,
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Department of Biology, Duke University, Durham, North Carolina, United States of America,
Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts, United States of America
School of Biosciences, Hatherly Laboratories, University of Exeter, Exeter, United Kingdom
School of Biological Sciences, The University of Texas at Austin, Austin, Texas, United States of America
Department of Biology, Duke University, Durham, North Carolina, United States of America
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