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Contributed equally to this work with:
Current address: Department of Brain and Cognitive Sciences, Picower Institute for Learning and Memory, Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America
Department of Cell and Molecular Biology, Karolinska Institute, Stockholm, Sweden
Current address: Medical Research Council (MRC) Laboratory of Molecular Biology, Cambridge, United Kingdom
Department of Neuroscience, Karolinska Institute, Stockholm, Sweden
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