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Contributed equally to this work with:
Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America
Current address: University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America
Department of Brain and Cognitive Sciences, Massachusetts Institute of Technology, Boston, Massachusetts, United States of America
Current address: Crick-Jacobs Center for Computational and Theoretical Biology, Salk Institute, La Jolla, California, United States of America
Department of Biology, Massachusetts Institute of Technology, Boston, Massachusetts, United States of America
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