About the Authors

Ziyue Gao

ziyuegao@stanford.edu (ZG); mp3284@columbia.edu (MP)

Current address: Howard Hughes Medical Institute, Stanford University, Stanford, California, United States of America

Affiliation Committee on Genetics, Genomics and Systems Biology, University of Chicago, Chicago, Illinois, United States of America

Minyoung J. Wyman

Affiliation Department of Biological Sciences, Columbia University, New York, New York, United States of America

Guy Sella

Affiliation Department of Biological Sciences, Columbia University, New York, New York, United States of America

Molly Przeworski

ziyuegao@stanford.edu (ZG); mp3284@columbia.edu (MP)

Affiliations Department of Biological Sciences, Columbia University, New York, New York, United States of America, Department of Systems Biology, Columbia University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZG GS MP. Performed the experiments: ZG MJW. Analyzed the data: ZG MJW. Wrote the paper: ZG GS MP.