About the Authors

Ileana Soto

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Leah C. Graham

Affiliations The Jackson Laboratory, Bar Harbor, Maine, United States of America, Sackler School of Graduate Biomedical Sciences, Tufts University, Boston, Massachusetts, United States of America

Hannah J. Richter

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Stephen N. Simeone

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Jake E. Radell

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Weronika Grabowska

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

W. Keith Funkhouser

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Megan C. Howell

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Gareth R. Howell

gareth.howell@jax.org

Affiliations The Jackson Laboratory, Bar Harbor, Maine, United States of America, Sackler School of Graduate Biomedical Sciences, Tufts University, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GRH IS. Performed the experiments: IS LCG HJR SNS JER WG WKF MCH. Analyzed the data: IS GRH. Wrote the paper: IS GRH.