About the Authors

Daniel P. Beiting

Affiliation Department of Pathobiology, University of Pennsylvania, Philadelphia, Pennsylvania, Philadelphia, United States of America

Shinya Hidano

Affiliation Department of Pathobiology, University of Pennsylvania, Philadelphia, Pennsylvania, Philadelphia, United States of America

Julie E. Baggs

Affiliation Department of Pharmacology and the Institute for Translational Medicine and Therapeutics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jeanne M. Geskes

Affiliation Department of Pharmacology and the Institute for Translational Medicine and Therapeutics, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Qun Fang

Affiliation Department of Pathobiology, University of Pennsylvania, Philadelphia, Pennsylvania, Philadelphia, United States of America

E. John Wherry

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Christopher A. Hunter

Affiliation Department of Pathobiology, University of Pennsylvania, Philadelphia, Pennsylvania, Philadelphia, United States of America

David S. Roos

droos@sas.upenn.edu (DSR); cherrys@mail.med.upenn.edu (SC)

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sara Cherry

droos@sas.upenn.edu (DSR); cherrys@mail.med.upenn.edu (SC)

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DPB SC DSR JEB JMG CAH. Performed the experiments: DPB SH QF. Analyzed the data: DPB SC CAH. Contributed reagents/materials/analysis tools: EJW. Wrote the paper: DPB SC.