About the Authors

Takashi Nobusawa

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Yoko Okushima

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Noriko Nagata

Affiliation Faculty of Science, Japan Women's University, Bunkyo-ku, Tokyo, Japan

Mikiko Kojima

Affiliation RIKEN Plant Science Center, Tsurumi, Yokohama, Japan

Hitoshi Sakakibara

Affiliation RIKEN Plant Science Center, Tsurumi, Yokohama, Japan

Masaaki Umeda

mumeda@bs.naist.jp

Affiliations Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan, JST, CREST, Ikoma, Nara, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Prepared data for submission to databases and did the submission: TN. Conceived and designed the experiments: TN YO HS MU. Performed the experiments: TN NN MK. Analyzed the data: TN HS. Contributed reagents/materials/analysis tools: NN MK HS. Wrote the paper: TN YO MU.