About the Authors

Yulei Shang

Affiliations Department of Pathology, University of California San Francisco, San Francisco, California, United States of America, Pathology Service 113B, VA Medical Center, San Francisco, California, United States of America

Christina N. Doan

Affiliations Department of Pathology, University of California San Francisco, San Francisco, California, United States of America, Pathology Service 113B, VA Medical Center, San Francisco, California, United States of America

Thomas D. Arnold

Affiliations Department of Physiology, University of California San Francisco, San Francisco, California, United States of America, Department of Pediatrics, University of California San Francisco, San Francisco, California, United States of America

Sebum Lee

Affiliations Department of Pathology, University of California San Francisco, San Francisco, California, United States of America, Pathology Service 113B, VA Medical Center, San Francisco, California, United States of America

Amy A. Tang

Affiliations Department of Pathology, University of California San Francisco, San Francisco, California, United States of America, Pathology Service 113B, VA Medical Center, San Francisco, California, United States of America

Louis F. Reichardt

Affiliation Department of Physiology, University of California San Francisco, San Francisco, California, United States of America

Eric J. Huang

eric.huang2@ucsf.edu

Affiliations Department of Pathology, University of California San Francisco, San Francisco, California, United States of America, Pathology Service 113B, VA Medical Center, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: YS EJH. Performed the experiments: YS CND TDA SL AAT. Analyzed the data: YS CND EJH. Contributed reagents/materials/analysis tools: TDA SL AAT LFR. Wrote the paper: YS EJH.