About the Authors

Naoki Shinohara

Address Current address: Department of Developmental Biology and Neurosciences, Graduate School of Life Sciences, Tohoku University, Sendai, Japan

Affiliation Department of Biology, University of York, Heslington, York, United Kingdom

Catherine Taylor

Affiliations Department of Biology, University of York, Heslington, York, United Kingdom, Sainsbury Laboratory, University of Cambridge, Cambridge, United Kingdom

Ottoline Leyser

ol235@cam.ac.uk

Affiliations Department of Biology, University of York, Heslington, York, United Kingdom, Sainsbury Laboratory, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: OL NS CT. Performed the experiments: NS CT. Analyzed the data: OL NS CT. Contributed reagents/materials/analysis tools: OL NS CT. Wrote the paper: OL NS CT.