About the Authors

Georgios Trichas

Contributed equally to this work with: Georgios Trichas, Aaron M. Smith

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Aaron M. Smith

Contributed equally to this work with: Georgios Trichas, Aaron M. Smith

Affiliation Centre for Mathematical Biology, Mathematical Institute, University of Oxford, Oxford, United Kingdom

Natalia White

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Vivienne Wilkins

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Tomoko Watanabe

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Abigail Moore

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Bradley Joyce

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Jacintha Sugnaseelan

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Tristan A. Rodriguez

Affiliation Imperial College London, London, United Kingdom

David Kay

Affiliation Oxford University Computing Laboratory, Oxford, United Kingdom

Ruth E. Baker

Affiliation Centre for Mathematical Biology, Mathematical Institute, University of Oxford, Oxford, United Kingdom

Philip K. Maini

Affiliations Centre for Mathematical Biology, Mathematical Institute, University of Oxford, Oxford, United Kingdom, Oxford Centre for Integrative Systems Biology, Department of Biochemistry, Oxford, United Kingdom

Shankar Srinivas

shankar.srinivas@dpag.ox.ac.uk

Affiliation Department of Physiology Anatomy & Genetics, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: SS PKM GT AS NW. Performed the experiments: GT AS NW VW BJ JS. Analyzed the data: GT AS NW TW AM BJ DK REB PKM SS. Contributed reagents/materials/analysis tools: TAR. Wrote the paper: SS AS PM GT.