About the Authors

Nadia Litterman

Affiliations Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Neuroscience, Harvard Medical School, Boston, Massachusetts, United States of America

Yoshiho Ikeuchi

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Gilbert Gallardo

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Brenda C. O'Connell

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Mathew E. Sowa

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Steven P. Gygi

Affiliation Department of Cell Biology, Harvard Medical School, Boston, Massachusetts, United States of America

J. Wade Harper

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Azad Bonni

azad_bonni@hms.harvard.edu

Affiliations Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Neuroscience, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: NL AB. Performed the experiments: NL YI GG BCO. Analyzed the data: NL GG BCO. Contributed reagents/materials/analysis tools: NL MES JWH SPG. Wrote the paper: NL AB.