About the Authors

Shawn C. Little

Contributed equally to this work with: Shawn C. Little, Gašper Tkačik

Affiliation Howard Hughes Medical Institute, Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Gašper Tkačik

Contributed equally to this work with: Shawn C. Little, Gašper Tkačik

Affiliation Department of Physics and Astronomy, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Thomas B. Kneeland

Affiliation Joseph Henry Laboratories of Physics, Princeton University, Princeton, New Jersey, United States of America

Eric F. Wieschaus

Affiliation Howard Hughes Medical Institute, Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Thomas Gregor

tg2@princeton.edu

Affiliations Joseph Henry Laboratories of Physics, Princeton University, Princeton, New Jersey, United States of America, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: SCL GT TG. Performed the experiments: SCL TG. Analyzed the data: SCL GT TBK TG. Contributed reagents/materials/analysis tools: SCL GT TBK EFW TG. Wrote the paper: SCL GT EFW TG.