About the Authors

Yingrui Li

Contributed equally to this work with: Yingrui Li, Jingde Zhu, Geng Tian, Ning Li, Qibin Li

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Jingde Zhu

Contributed equally to this work with: Yingrui Li, Jingde Zhu, Geng Tian, Ning Li, Qibin Li

Affiliations Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China, Epigentic Laboratory, Bio-X Center, Shanghai JiaoTong University, Shanghai, China

Geng Tian

Contributed equally to this work with: Yingrui Li, Jingde Zhu, Geng Tian, Ning Li, Qibin Li

Affiliations Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, China, The Graduate University of Chinese Academy of Sciences, Beijing, China, BGI-Shenzhen, Shenzhen, Guangdong, China

Ning Li

Contributed equally to this work with: Yingrui Li, Jingde Zhu, Geng Tian, Ning Li, Qibin Li

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Qibin Li

Contributed equally to this work with: Yingrui Li, Jingde Zhu, Geng Tian, Ning Li, Qibin Li

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Mingzhi Ye

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Hancheng Zheng

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Jian Yu

Affiliation Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China

Honglong Wu

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Jihua Sun

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Hongyu Zhang

Affiliation Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China

Quan Chen

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Ruibang Luo

Affiliations BGI-Shenzhen, Shenzhen, Guangdong, China, School of Bioscience and Biotechnology, South China University of Technology, Guangzhou, China

Minfeng Chen

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Yinghua He

Affiliation Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China

Xin Jin

Affiliations BGI-Shenzhen, Shenzhen, Guangdong, China, School of Bioscience and Biotechnology, South China University of Technology, Guangzhou, China

Qinghui Zhang

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Chang Yu

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Guangyu Zhou

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Jinfeng Sun

Affiliation Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China

Yebo Huang

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Huisong Zheng

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Hongzhi Cao

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Xiaoyu Zhou

Affiliation Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China

Shicheng Guo

Affiliation Cancer Epigenetics and Gene Therapy Program, Shanghai Cancer Institute, Shanghai Jiaotong University, Shanghai, China

Xueda Hu

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Xin Li

Affiliation Kunming Institute of Zoology, Chinese Academy of Sciences, Yunnan, China

Karsten Kristiansen

Affiliation Department of Biology, University of Copenhagen, Copenhagen, Denmark

Lars Bolund

Affiliations BGI-Shenzhen, Shenzhen, Guangdong, China, Institute of Human Genetics, University of Aarhus, Aarhus, Denmark

Jiujin Xu

Affiliation Institute of Genetics and Developmental Biology, Chinese Academy of Human Sciences, Beijing, China

Wen Wang

Affiliation Kunming Institute of Zoology, Chinese Academy of Sciences, Yunnan, China

Huanming Yang

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Jian Wang

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Ruiqiang Li

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Stephan Beck

zhangxq@genomics.org.cn (XZ); wangj@genomics.org.cn (JW); s.beck@ucl.ac.uk (SB)

Affiliation UCL Cancer Institute, University College London, London, United Kingdom

Jun Wang

zhangxq@genomics.org.cn (XZ); wangj@genomics.org.cn (JW); s.beck@ucl.ac.uk (SB)

Affiliations BGI-Shenzhen, Shenzhen, Guangdong, China, Department of Biology, University of Copenhagen, Copenhagen, Denmark

Xiuqing Zhang

zhangxq@genomics.org.cn (XZ); wangj@genomics.org.cn (JW); s.beck@ucl.ac.uk (SB)

Affiliation BGI-Shenzhen, Shenzhen, Guangdong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: JZ GT WW HY JW SB JW XZ. Performed the experiments: JZ GT MY JY JS HZ MC YH QZ JS YH HZ XZ SG XH LB JX XZ. Analyzed the data: YL NL QL HZ HW RL XJ GZ HC KK RL SB JW. Contributed reagents/materials/analysis tools: YL JZ NL QL HZ QC CY RL XZ. Wrote the paper: YL XL SB JW.