About the Authors

Caroline H. Ko

Contributed equally to this work with: Caroline H. Ko, Yujiro R. Yamada, David K. Welsh

Address Current address: Department of Chemistry, Northwestern University, Evanston, Illinois, United States of America

Affiliations Department of Neuroscience, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America, Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America, Department of Psychology, University of Toronto, Toronto, Ontario, Canada

Yujiro R. Yamada

Contributed equally to this work with: Caroline H. Ko, Yujiro R. Yamada, David K. Welsh

Affiliation Department of Mathematics, University of Michigan, Ann Arbor, Michigan, United States of America

David K. Welsh

Contributed equally to this work with: Caroline H. Ko, Yujiro R. Yamada, David K. Welsh

Affiliations Department of Cell and Developmental Biology, University of California, San Diego, La Jolla, California, United States of America, Department of Psychiatry, University of California, San Diego, La Jolla, California, United States of America, Veterans Affairs San Diego Healthcare System, San Diego, California, United States of America

Ethan D. Buhr

Address Current address: Department of Ophthalmology, University of Washington, Seattle, Washington, United States of America

Affiliation Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America

Andrew C. Liu

Address Current address: Department of Biology, University of Memphis, Memphis, Tennessee, United States of America

Affiliation Genomics Institute of Novartis Research Foundation, San Diego, California, United States of America

Eric E. Zhang

Affiliation Genomics Institute of Novartis Research Foundation, San Diego, California, United States of America

Martin R. Ralph

Affiliations Department of Psychology, University of Toronto, Toronto, Ontario, Canada, Center for Biological Timing and Cognition, University of Toronto, Toronto, Ontario, Canada

Steve A. Kay

Affiliation Department of Cell and Developmental Biology, University of California, San Diego, La Jolla, California, United States of America

Daniel B. Forger

Affiliations Department of Mathematics, University of Michigan, Ann Arbor, Michigan, United States of America, Center for Computational Medicine and Bioinformatics, University of Michigan, Ann Arbor, Michigan, United States of America

Joseph S. Takahashi

Joseph.Takahashi@UTSouthwestern.edu

Affiliations Department of Neuroscience, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America, Department of Neurobiology and Physiology, Northwestern University, Evanston, Illinois, United States of America, Howard Hughes Medical Institute, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: CHK YRY DKW SAK DBF JST. Performed the experiments: CHK YRY DKW EDB ACL EEZ DBF. Analyzed the data: CHK YRY DKW EDB ACL EEZ DBF JST. Contributed reagents/materials/analysis tools: MRR SAK. Wrote the paper: CHK YRY DBF JST.