About the Authors

Yong E. Zhang

Affiliation Department of Ecology and Evolution, the University of Chicago, Chicago, Illinois, United States of America

Maria D. Vibranovski

Affiliation Department of Ecology and Evolution, the University of Chicago, Chicago, Illinois, United States of America

Patrick Landback

Affiliation Department of Ecology and Evolution, the University of Chicago, Chicago, Illinois, United States of America

Gabriel A. B. Marais

Affiliation Université de Lyon, Centre National de la Recherche Scientifique, Laboratoire de Biométrie et Biologie évolutive, Villeurbanne, France

Manyuan Long

mlong@uchicago.edu

Affiliation Department of Ecology and Evolution, the University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: YEZ MDV GABM ML. Performed the experiments: YEZ GABM. Analyzed the data: YEZ MDV PL GABM ML. Contributed reagents/materials/analysis tools: YEZ. Wrote the paper: YEZ MDV PL GABM ML.