About the Authors

Yu Zhang

Affiliation Laboratory of Cellular and Developmental Biology, National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

John H. Malone

Affiliation Laboratory of Cellular and Developmental Biology, National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Sara K. Powell

Affiliation Department of Pharmacology and Cancer Biology, Duke University Medical Center, Durham, North Carolina, United States of America

Vipul Periwal

Affiliation Laboratory of Biological Modeling, National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Eric Spana

Affiliation Department of Biology, Duke University, Durham, North Carolina, United States of America

David M. MacAlpine

Affiliation Department of Pharmacology and Cancer Biology, Duke University Medical Center, Durham, North Carolina, United States of America

Brian Oliver

oliver@helix.nih.gov

Affiliation Laboratory of Cellular and Developmental Biology, National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: YZ SKP ES DMM BO. Performed the experiments: YZ SKP. Analyzed the data: YZ JHM SKP VP DMM BO. Contributed reagents/materials/analysis tools: JHM ES. Wrote the paper: YZ BO.