About the Authors

Debora Bogani

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Pam Siggers

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Rachel Brixey

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Nick Warr

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Sarah Beddow

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Jessica Edwards

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Debbie Williams

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Dagmar Wilhelm

Affiliation The Institute of Molecular Bioscience, University of Queensland, Brisbane, Australia

Peter Koopman

Affiliation The Institute of Molecular Bioscience, University of Queensland, Brisbane, Australia

Richard A. Flavell

Affiliation Department of Immunobiology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Hongbo Chi

Affiliation Department of Immunology, St. Jude Children's Research Hospital, Memphis, Tennessee, United States of America

Harry Ostrer

Affiliation Human Genetics Program, New York University School of Medicine, New York, New York, United States of America

Sara Wells

Affiliation The Mary Lyon Centre, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Michael Cheeseman

Affiliations Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom, The Mary Lyon Centre, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Andy Greenfield

a.greenfield@har.mrc.ac.uk

Affiliation Mammalian Genetics Unit, Medical Research Council (MRC) Harwell, Oxfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: DB PS RB NW DW MC AG. Performed the experiments: DB PS RB NW SB JE DW SW MC. Analyzed the data: DB PS RB NW DW MC AG. Contributed reagents/materials/analysis tools: DW PK RAF HC HO. Wrote the paper: AG.