About the Authors

Graham E Little

Contributed equally to this work with: Graham E Little, Guillermina López-Bendito

Affiliation Smurfit Institute of Genetics, Trinity College Dublin, Dublin, Ireland

Guillermina López-Bendito

Contributed equally to this work with: Graham E Little, Guillermina López-Bendito

To whom correspondence should be addressed. E-mail: g.lbendito@umh.es (GL-B); zoltan.molnar@dpag.ox.ac.uk (ZM); Kevin.Mitchell@tcd.ie (KJM)

Affiliations Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford, Oxfordshire, United Kingdom , Instituto de Neurociencias de Alicante, CSIC & Universidad Miguel Hernández, San't Joan d'Alacant, Spain

Annette E Rünker

Affiliation Smurfit Institute of Genetics, Trinity College Dublin, Dublin, Ireland

Noelia García

Affiliation Instituto de Neurociencias de Alicante, CSIC & Universidad Miguel Hernández, San't Joan d'Alacant, Spain

Maria C Piñon

Affiliation Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Alain Chédotal

Affiliations INSERM, UMR S968, Institut de la Vision, Department of Development, Paris, France , UPMC Univ Paris 06, UMR S968, Institut de la Vision, Paris, France

Zoltán Molnár

To whom correspondence should be addressed. E-mail: g.lbendito@umh.es (GL-B); zoltan.molnar@dpag.ox.ac.uk (ZM); Kevin.Mitchell@tcd.ie (KJM)

Affiliation Department of Physiology, Anatomy and Genetics, University of Oxford, Oxford, Oxfordshire, United Kingdom

Kevin J Mitchell

To whom correspondence should be addressed. E-mail: g.lbendito@umh.es (GL-B); zoltan.molnar@dpag.ox.ac.uk (ZM); Kevin.Mitchell@tcd.ie (KJM)

Affiliation Smurfit Institute of Genetics, Trinity College Dublin, Dublin, Ireland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

GEL, GL-B, AER, ZM, and KJM conceived and designed the experiments. GEL, GL-B, AER, NG, MCP, AC, and ZM performed the experiments. GEL, GL-B, AER, NG, MCP, ZM, and KJM analyzed the data. AC contributed reagents/materials/analysis tools. GEL, GL-B, ZM, and KJM wrote the paper.