About the Authors

Erin K. McCarthy Campbell

Affiliation Department of Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Adam D Werts

Affiliation Department of Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Bob Goldstein

Affiliation Department of Biology, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

EKMC and BG conceived and designed the experiments. EKMC and ADW performed the experiments. EKMC, ADW, and BG analyzed the data. EKMC and BG wrote the paper.