About the Authors

Peter Serrano

Contributed equally to this work with: Peter Serrano, Eugenia L Friedman

Address ¤ Current address: Department of Psychology and the Gene Research Center, Hunter College, City University of New York, New York, New York, United States of America

Affiliation Department of Physiology and Pharmacology, The Robert F. Furchgott Center for Neural and Behavioral Science, State University of New York, Brooklyn, New York, United States of America

Eugenia L Friedman

Contributed equally to this work with: Peter Serrano, Eugenia L Friedman

Affiliations Department of Psychiatry, University of Wisconsin-Madison Medical School, Madison, Wisconsin, United States of America , Neuroscience Training Program, University of Wisconsin-Madison Medical School, Madison, Wisconsin, United States of America

Jana Kenney

Affiliation Institute of Physiology, Academy of Sciences of the Czech Republic, Prague, Czech Republic

Stephen M Taubenfeld

Affiliation Fishberg Department of Neuroscience, Mount Sinai School of Medicine, Icahn Medical Institute, New York, New York, United States of America

Joshua M Zimmerman

Affiliation Neuroscience Program, University of Michigan, Ann Arbor, Michigan, United States of America

John Hanna

Affiliation Department of Physiology and Pharmacology, The Robert F. Furchgott Center for Neural and Behavioral Science, State University of New York, Brooklyn, New York, United States of America

Cristina Alberini

Affiliation Fishberg Department of Neuroscience, Mount Sinai School of Medicine, Icahn Medical Institute, New York, New York, United States of America

Ann E Kelley

† Deceased.

Affiliation Department of Psychiatry, University of Wisconsin-Madison Medical School, Madison, Wisconsin, United States of America

Stephen Maren

Affiliations Neuroscience Program, University of Michigan, Ann Arbor, Michigan, United States of America , Department of Psychology, University of Michigan, Ann Arbor, Michigan, United States of America

Jerry W Rudy

Affiliation Department of Psychology, Center for Neuroscience, University of Colorado, Boulder, Colorado, United States of America

Jerry C. P Yin

Affiliations Department of Psychiatry, University of Wisconsin-Madison Medical School, Madison, Wisconsin, United States of America , Department of Genetics, University of Wisconsin-Madison Medical School, Madison, Wisconsin, United States of America

Todd C Sacktor

To whom correspondence should be addressed. E-mail: tsacktor@downstate.edu (TCS); afenton@downstate.edu (AAF)

Affiliations Department of Physiology and Pharmacology, The Robert F. Furchgott Center for Neural and Behavioral Science, State University of New York, Brooklyn, New York, United States of America , Department of Neurology, The Robert F. Furchgott Center for Neural and Behavioral Science, State University of New York, Brooklyn, New York, United States of America

André A Fenton

To whom correspondence should be addressed. E-mail: tsacktor@downstate.edu (TCS); afenton@downstate.edu (AAF)

Affiliation Department of Physiology and Pharmacology, The Robert F. Furchgott Center for Neural and Behavioral Science, State University of New York, Brooklyn, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PS, ELF, JK, SMT, and JMZ performed and analyzed the eight-arm radial maze, tone-fear, water maze, inhibitory avoidance, and contextual fear experiments, respectively. JH assisted with the eight-arm radial maze training. JWR performed the dorsal and ventral hippocampi contextual fear experiments. CA, AEK, SM, JWR, JCPY, TCS, and AAF supervised and designed the research. TCS and AAF wrote the manuscript.