About the Authors

Guo-Xiang Ruan

Affiliation Department of Biological Sciences, Vanderbilt University, Nashville, Tennessee, United States of America

Gregg C Allen

Affiliation Department of Biological Sciences, Vanderbilt University, Nashville, Tennessee, United States of America

Shin Yamazaki

Affiliation Department of Biological Sciences, Vanderbilt University, Nashville, Tennessee, United States of America

Douglas G McMahon

To whom correspondence should be addressed. E-mail: douglas.g.mcmahon@vanderbilt.edu

Affiliation Department of Biological Sciences, Vanderbilt University, Nashville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

G-XR and DGM conceived and designed the experiments. G-XR and GCA performed the experiments. G-XR analyzed the data. G-XR and SY contributed reagents/materials/analysis tools. G-XR and DGM wrote the paper.