About the Authors

Oi Kwan Wong

Affiliation Department of Biochemistry, Brandeis University, Waltham, Massachusetts, United States of America

Martin Guthold

Affiliation Department of Physics, Wake Forest University, Winston-Salem, North Carolina, United States of America

Dorothy A Erie

Affiliation Department of Chemistry and Curriculum Applied and Materials Sciences, University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Jeff Gelles

To whom correspondence should be addressed. E-mail: gelles@brandeis.edu

Affiliation Department of Biochemistry, Brandeis University, Waltham, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

OKW, MG, DAE, and JG conceived and designed the experiments. OKW and MG performed the experiments. OKW, MG, DAE, and JG analyzed the data. OKW, DAE, and JG wrote the paper. All authors reviewed and edited the manuscript.