About the Authors

Cathy Haag-Liautard

Address ¤ Current address: University of Fribourg, Institute of Ecology and Evolution, Fribourg, Switzerland

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Nicole Coffey

Affiliation Department of Biology, Indiana University, Bloomington, Indiana, United States of America

David Houle

Affiliation Department of Biological Science, Florida State University, Tallahassee, Florida, United States of America

Michael Lynch

Affiliation Department of Biology, Indiana University, Bloomington, Indiana, United States of America

Brian Charlesworth

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Peter D Keightley

To whom correspondence should be addressed. E-mail: keightley.pb2008@gmail.com

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DH was responsible for producing the Florida MA lines CH-L carried out the DHPLC analysis of MA lines, associated DNA sequencing, and pyrosequencing. NC and ML were responsible the direct sequencing of MA lines. PDK and CH-L analysed the data, with advice from BC. PDK drafted the paper. All authors revised the manuscript.