About the Authors

Esteban O Mazzoni

Address Current address: Pathology Department, Columbia University, New York, New York, United States of America

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Arzu Celik

Address Current address: Department of Molecular Biology and Genetics, Bogazici University, Istanbul, Turkey

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Mathias F Wernet

Address Current address: Department of Neurobiology, Stanford University, Stanford, California, United States of America

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Daniel Vasiliauskas

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Robert J Johnston

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Tiffany A Cook

Address Current address: Developmental Biology/Pediatric Ophthalmology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Franck Pichaud

Address Current address: Department of Anatomy and Developmental Biology, Medical Research Council Laboratory for Molecular Cell Biology and Cell Biology Unit, University College London, London, United Kingdom

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Claude Desplan

To whom correspondence should be addressed. E-mail: cd38@nyu.edu

Affiliation Center for Developmental Genetics, Department of Biology, New York University, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

EOM, AC, MFW, and CD conceived and designed the experiments. EOM, AC, MFW, and DV performed and analyzed the experiments. RJJ and TAC contributed reagents. TAC and FP made the initial observations. EOM, AC, and CD wrote the paper.