About the Authors

Karen R Lips, Michael W Sears
Department of Zoology, Southern Illinois University, Carbondale, Illinois, United States of America

Jay Diffendorfer
Illinois Natural History Survey, Champaign, Illinois, United States of America

Joseph R Mendelson III
Zoo Atlanta, Atlanta, Georgia, United States of America

Corresponding Author

klips@zoology.siu.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

All four coauthors—KRL, JD, JRM, and MWS—participated in the conception and design of the research, data collection and analyses, and writing of the paper.