About the Authors

Nelson S Yee

Contributed equally to this work with: Nelson S Yee, Weilong Gong

Address Current address: Department of Internal Medicine, University of Iowa, Carver College of Medicine, Iowa City, Iowa, United States of America

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Weilong Gong

Contributed equally to this work with: Nelson S Yee, Weilong Gong

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Ying Huang

Address Current address: College of Life Sciences, Nanjing Normal University, Nanjing, Peoples Republic of China

Affiliation National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland, United States of America

Kristin Lorent

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Amy C Dolan

Affiliation Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Richard J Maraia

Affiliation National Institute of Child Health and Human Development, National Institutes of Health, Bethesda, Maryland, United States of America

Michael Pack

To whom correspondence should be addressed. E-mail: mpack@mail.med.upenn.edu

Affiliations Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America , Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

NSY, WG, YH, KL, RJM and MP conceived and designed the experiments and analyzed the data. NSY, WG, YH, KL, and ACD performed the experiments. RJM and MP wrote the paper.