About the Authors

HoJung Cho

Contributed equally to this work with: HoJung Cho, Henrik Jönsson

Affiliation Department of Biomedical Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America

Henrik Jönsson

Contributed equally to this work with: HoJung Cho, Henrik Jönsson

Affiliation Department of Theoretical Physics, Lund University, Lund, Sweden

Kyle Campbell

Affiliation Department of Physics, University of California San Diego, La Jolla, California, United States of America

Pontus Melke

Affiliation Department of Theoretical Physics, Lund University, Lund, Sweden

Joshua W Williams

Affiliation Department of Biological Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Bruno Jedynak

Affiliation Center for Imaging Science, The Johns Hopkins University, Baltimore, Maryland, United States of America

Ann M Stevens

Affiliation Department of Biological Sciences, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, United States of America

Alex Groisman

To whom correspondence should be addressed. E-mail: alev@jhu.edu (AL), agroisman@ucsd.edu (AG)

Affiliation Department of Physics, University of California San Diego, La Jolla, California, United States of America

Andre Levchenko

To whom correspondence should be addressed. E-mail: alev@jhu.edu (AL), agroisman@ucsd.edu (AG)

Affiliation Department of Biomedical Engineering, The Johns Hopkins University, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HC, HJ, AG, and AL designed research. HC, HJ, PM, and AL performed the experiments and simulations. KC, AL and AG designed and fabricated the microfluidic devices. JWW, BJ, and AMS contributed new reagents/analysis tools. HC, HJ, PM, and AL analyzed data. HC, HJ, PM, AG, and AL wrote the paper.