About the Authors

Nadin Rohland

Contributed equally to this work with: Nadin Rohland, Anna-Sapfo Malaspinas

Affiliation Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany

Anna-Sapfo Malaspinas

Contributed equally to this work with: Nadin Rohland, Anna-Sapfo Malaspinas

Affiliations Department of Integrative Biology, University of California Berkeley, Berkeley, California, United States of America , Department of Zoology and Animal Biology, University of Geneva, Geneva, Switzerland

Joshua L Pollack

Affiliation Department of Integrative Biology, University of California Berkeley, Berkeley, California, United States of America

Montgomery Slatkin

Affiliation Department of Integrative Biology, University of California Berkeley, Berkeley, California, United States of America

Paul Matheus

Affiliation Alaska Quaternary Center, University of Alaska Fairbanks, Fairbanks, Alaska, United States of America

Michael Hofreiter

To whom correspondence should be addressed. E-mail: hofreite@eva.mpg.de

Affiliation Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

NR and MH conceived and designed the experiments. NR performed the experiments. PM collected data. ASM, JLP, and MS analyzed the data. NR, ASM, MS, PM, and MH contributed to writing the paper.