About the Authors

Stuart Humphries

To whom correspondence should be addressed. E-mail: s.humphries@sheffield.ac.uk

¤ Current address: Department of Animal and Plant Sciences, University of Sheffield, Sheffield, United Kingdom

Affiliation School of Biological Sciences, University of Reading, Reading, United Kingdom

Richard H. C Bonser

Affiliation School of Construction Management and Engineering, University of Reading, Reading, United Kingdom

Mark P Witton

Affiliation School of Earth and Environmental Sciences, University of Portsmouth, Portsmouth, United Kingdom

David M Martill

Affiliation School of Earth and Environmental Sciences, University of Portsmouth, Portsmouth, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SH conceived and designed the experiments. All authors performed the experiments and wrote the paper. SH and RHCB analyzed the data. SH, RHCB, and DMM contributed reagents/materials/analysis tools.