About the Authors

Xiaoyu Zhang

Contributed equally to this work with: Xiaoyu Zhang, Oliver Clarenz

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Oliver Clarenz

Contributed equally to this work with: Xiaoyu Zhang, Oliver Clarenz

Affiliations Institute of Molecular Plant Science, University of Edinburgh, Edinburgh, United Kingdom , School of Biology, University of Edinburgh, Edinburgh, United Kingdom

Shawn Cokus

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Yana V Bernatavichute

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America , Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America

Matteo Pellegrini

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Justin Goodrich

Affiliations Institute of Molecular Plant Science, University of Edinburgh, Edinburgh, United Kingdom , School of Biology, University of Edinburgh, Edinburgh, United Kingdom

Steven E Jacobsen

To whom correspondence should be addressed. E-mail: Jacobsen@ucla.edu

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America , Howard Hughes Medical Institute, University of California Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JG and SEJ conceived and designed the experiments and edited the paper. XZ, OC, and YVB performed the experiments. XZ, SC, and MP analyzed the data. XZ wrote the paper.