About the Authors

Takashi Matsuo

To whom correspondence should be addressed. E-mail: mts@comp.metro-u.ac.jp

Affiliation Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo, Japan

Shigeru Sugaya

Address ¤ Current address: School of Medicine, Chiba University, Chiba-shi, Chiba, Japan

Affiliation Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo, Japan

Jyunichiro Yasukawa

Affiliation Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo, Japan

Toshiro Aigaki

Affiliation Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo, Japan

Yoshiaki Fuyama

Affiliation Department of Biological Sciences, Tokyo Metropolitan University, Hachioji, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

TM and YF conceived and designed the experiments. TM, SS, and JY performed the experiments and analyzed the data. TA contributed reagents/materials/analysis tools. TM wrote the paper.